Delphacid predators and parasites are of particular interest to find out what natural enemies might be used to control delphacids and to better understand planthopper population dynamics. The most important predators of delphacids include (but is not limited to) spiders, ground beetles, lady beetles, predaceous mirids, damsel bugs and other generalized predators.  Perhaps spiders are the most important, but mirids (Tytthus spp.) have been shown to be very important in some settings.  Egg predators include microhymenopteran wasps such as Encyrtidae, Eulophidae and Mymaridae.  The most important parasites are Strepsiptera (Elenchidae and Halictophagidae), dryinid wasps, and big-headed flies (Pipunculidae). Dryinid and strepsipteran parasites are often seen in museum specimens and in field populations.

Caenodelphax atridorsum female with a parasite (Dryinidae)

Ricehoppers.net has compiled this working summary of parasites of rice pests (here for PDF) and here: http://ricehoppers.net/wp-content/uploads/2010/03/interim-report-ii-appendix-1-gm-gurr.pdf.

Recently, Dr. Eduardo Virla and Dra. Erica Luft of the Laboratorio de Enemigos Naturales de Plagas Agrícolas del NOA at the División Control Biológico de Plagas, CONICET, Tucumán, Argentina (http://www.biocontroltucuman.com.ar/) provided me with a more complete list of delphacid parasites.  Dr. Virla has a webpage on Neotropical Dryinidae (www.parasitoides.com.ar) (as of this writing it is down because of server problems).  The list was send with links to the dryinid species pages, which I will reinsert at next opportunity.  It is with immense gratitude for their work that I provide this parasite list and references below.

Dryinidae
Delphacid taxon	Country: Dryinid species	Reference
Delphacidae, unknown species	Argentina: Haplogonatopus hernandezae Olmi 1984	Virla & Olmi 1998b
	Ecuador: Gonatopus flavoides (Olmi 1984)	Olmi 1987b
Anchidelphax albidens (Crawford)	Ecuador: Gonatopus vignai (Olmi 1987)	Guglielmino & Olmi 1997
Calligypona sp.	Dutch Antillies: Gonatopus cobbeni (Olmi 1987)	Olmi 1987c
Delphacodes haywardi Muir	Argentina: Haplogonatopus hernandezae Olmi 1984	Virla & Olmi 1998b
	Argentina: Gonatopus chilensis (Olmi 1991)	Olmi & Virla 1993
Delphacodes kuscheli Fennah	Argentina: Haplogonatopus hernandezae Olmi 1984	Virla & Olmi 1998b
	Argentina: Gonatopus chilensis (Olmi 1991)	Olmi & Virla 1993
	Argentina: Gonatopus lauti Virla 1998	Virla, 2001
	Argentina: Gonatopus bonaerensis Virla 1997	Cuezzo & Virla, 2001; Virla, 2004
Delphacodes latus Remes Lenicov & Tesón	Argentina: Gonatopus bonaerensis Virla 1997	Cuezzo & Virla, 2001; Virla, 2004
Delphacodes nr albinotata (Crawford) (this species currently Delphacodes arcuata Beamer, 1948)	Ecuador: Gonatopus flavoides (Olmi 1984)	Guglielmino & Olmi 1997
Delphacodes sitarea Remes Lenicov & Tesón	Argentina: Haplogonatopus hernandezae Olmi 1984	Virla & Olmi 1998b
	Argentina: Gonatopus chilensis (Olmi 1991)	Virla 1995
	Argentina: Gonatopus bonaerensis Virla 1997	Cuezzo & Virla, 2001; Virla, 2004
Dicranotropis sp.	Argentina: Gonatopus bonaerensis Virla 1997	Cuezzo & Virla, 2001; Virla, 2004
Dicranotropis fuscoterminata (Berg)	Argentina: Gonatopus chilensis (Olmi 1991)	Virla 1995
Megamelus scutellaris (Berg)	Argentina: Gonatopus hilaris Olmi 1995	Sosa et al., 2005
Nesosydne alcmaeon Fennah	Ecuador: Haplogonatopus crucianus Olmi 1986	Abedrabbo et al., 1990
Nesosydne olipor Fennah	Ecuador: Haplogonatopus crucianus Olmi 1986	Abedrabbo et al., 1990
Peregrinus maidis (Ashmead)	Argentina: Haplogonatopus hernandezae Olmi 1984	Cuezzo & Virla, 2001
	Argentina: Gonatopus chilensis (Olmi 1991)	Cuezzo & Virla, 2001
	Argentina: Gonatopus bonaerensis Virla 1997	Cuezzo & Virla, 2001; Virla, 2004
Perkinsiella saccharicida Kirkaldy	Colombia: Gonatopus morenoi (Olmi 1984)	Olmi 1984
	Perú: Gonatopus krombeini Olmi 1998	Olmi 1989
Saccharosydne saccharivora (Westwood)	USA: Gonatopus arizonicus (Perkins 1907)	Olmi 1984
	Jamaica, Guadeloupe: Gonatopus variistriatus (Fenton 1927)	Olmi 1984; Olmi 1998a
	Bahamas: Gonatopus variistriatus (Fenton 1927)	Guglielmino & Olmi 1997
Sogata sp. (probably this refers to a taxon now in Sogatella or Tagosodes)	Cuba: Echthrodelphax krombeini Olmi 1984	Alayo 1974, Guglielmino & Olmi 1997
Sogatella sp.	Guadaloupe: Gonatopus cubensis (Richards, 1969)	According with Olmi (1984), probably an error, because this hopper belongs to Delphacidae
	Argentina: Gonatopus chilensis (Olmi 1991)	Virla 1995
Stobaera sp.	USA: Gonatopus arizonicus (Perkins 1907)	RCL Perkins 1907
Struebingianella lugubrina (Boheman)	Dutch Antilles: Haplogonatopus hernandezae Olmi 1984	Olmi 1987c
Tagosodes orizicolus (Muir)	Colombia: Gonatopus flavoides (Olmi 1984)	Guglielmino & Olmi 1997
	Colombia, Panamá, Costa Rica, Perú: Haplogonatopus hernandezae Olmi 1984	Olmi 1984; Guglielmino & Olmi 2006
	Perú: Gonatopus chilensis (Olmi 1991)	Guglielmino & Olmi 2006
	Colombia: Gonatopus morenoi (Olmi 1984)	Guglielmino & Olmi 1997
Toya argentinensis Remes Lenicov & Tesón (now Metadelphax argentinensis)	Argentina: Haplogonatopus hernandezae Olmi 1984	Virla & Olmi, 1998
	Argentina: Gonatopus chilensis (Olmi 1991)	Virla & Olmi, 1998
Toya propinqua (Fieber) (Now Metadelphax propinqua)	Argentina: Haplogonatopus hernandezae Olmi 1984	Virla & Olmi 1998b
	Argentina: Gonatopus chilensis (Olmi 1991)	Olmi & Virla 1993
	Argentina: Gonatopus lauti Virla 1998	Virla, 2001
	Argentina: Gonatopus bonaerensis Virla 1997	Cuezzo & Virla, 2001; Virla, 2004
	Argentina: Gonatopus flavoides (Olmi 1984)	Virla & Olmi 2007b
PIPUNCULIDAE (Diptera)
Delphacidae	Argentina: Cephalops penepauculus (Hardy)
Metadelphax propinqua (Fieber)		Virla & Rafael 1997
Dicranotropis fuscoterminata Berg

 

Egg Parasitods

Family	Genus	species	Delphacid host	Delphacid host	Locality	Reference
Eulophidae	Aprostocetus	infulatus	Peregrinus maidis	Corn	Argentina	Luft Albarracin & Triapitsyn, 2007
Mymaridae	Anagrus	breviphragma	Delphacodes kuscheli	Oat	Argentina	Triapitsyn 1997
Mymaridae	Anagrus	flaveolus	Peregrinus maidis	Corn	Argentina	Luft Albarracin et al. 2009
Mymaridae	Anagrus	flaveolus	Delphacodes kuscheli	Oat	Argentina	Luft Albarracin et al. 2009
Mymaridae	Anagrus	flaveolus	Saccharosydne subandina	Cortaderia	Argentina	Triapitsyn et al. 2011
Mymaridae	Anagrus	flaveolus	Metadelphax propinqua	Bermudagrass	Argentina	Luft Albarracin et al. 2009
Mymaridae	Anagrus	miriamae	Delphacodes sitarea	Grass	Argentina	Triapitsyn & Virla 2004

Reference list

Abedrabbo, S., J. Kathirithamby, and M. Olmi. 1990. Contribution to the knowledge of the Elenchidae (Strepsiptera) and Dryinidae (Hymenoptera: Chrysidoidea) of the Galapagos Islands. Bollettino dell'Istituto di Entomologia della Universita degli Studi di Bologna XLV (1990): 121–128.

Alayo, P. 1974. Introduccion al estudio de los himenopteros de Cuba. Superfamilia. Bethyloidea. Serie Biologica, Academia de Ciencias de Cuba, 52:1–24.

Cuezzo, F. and E. Virla. 2001. Interacción entre Delphacidae e Hymenoptera en cultivos de maíz en la Argentina. Revista de la Sociedad Entomológica Argentina 60 (1-4): 35-37. ISSN 0373-5680.

De Santis, L., Virla, E. and Maragliano, R. 1992. Presencia de Anagrus flaveolus en la República Argentina parasi¬toide de un insecto dañino del trigo y maíz (Insecta - Hymenoptera - Mymaridae). Revista de la Facultad de Agronomía, La Plata, 13(1): 19-23. ISSN 0325-9250.

Guglielmino A. and M. Olmi. 2006. A host-parasite catalog of world Dryinidae (Hymenoptera: Chrysidoidea): first supplement. Zootaxa 1139: 35-62.

Guglielmino A. and M. Olmi. 1997. A host-parasite catalog of world Dryinidae (Hymenoptera: Chrysidoidea). Contributions on Entomology, International 2: 165-298.

Guglielmino A. and M. Olmi. 2007. A host-parasite catalog of world Dryinidae (Hymenoptera: Chrysidoidea): second supplement. Bollettino di Zoologia Agraria e Bachicoltura (Ser. ii) 39: 121-129.

Laguna, I.; A. Avila; E. Virla; M. P. Giménez Pecci; A. M. de Remes Lenicov; R. Mariani; J. Garay and M. Fiorona. 2003. Difusión del Mal de Río Cuarto (MRCV) del maíz, su vector, delfácidos asociados, sus enemigos naturales y hospedantes alternativos en la región de Cuyo de Argentina. Brazilian Phytopathology 28(supl.): 239. ISSN 0100-4158.

Liljesthröm, G. G. and E. Virla. 2004. Density dependent parasitism of Delphacodes kuscheli (Homoptera: Dephacidae) eggs by Anagrus flaveolus (Hymenoptera: Mymaridae): Influence of egg patchiness and density. Biocontrol Science and Technology 14(2): 107-115. ISSN 0958-3157.

Luft Albarracin, E. and Triapitsyn, S. V. 2007. Aprostocetus (Ootetrastichus) infulatus (Hymenoptera: Eulophidae): description of the male, new distribution and first host records. Zootaxa 1438: 65 -68.

Luft Albarracin, E., S. Triapitsyn and E. Virla. 2009. An annotated key to the genera of Mymaridae (Hymenoptera: Chalcidoidea) in Argentina. Zootaxa 2129: 1-28. ISSN 1175-5326, 1175-5334 (on line).

Luft Albarracin, E.; Triapitsyn, S. V.; and Virla, E. G. 2009. Annotated key to the genera of Mymaridae (Hymenoptera: Chalcidoidea) in Argentina. Zootaxa 2129: 1-28.

Mangione, S. and E. Virla. 2004. Morfología de los estados preimaginales de Gonatopus bonaerensis, y consideraciones sobre la morfología interna de sus larvas inmaduras (Hymenoptera, Dryinidae) Acta Zoológica Lilloana 47(1-2): 91-102. ISSN 0065-1729.

Mariani, R., L. Vera and E. Virla. 1997. Aportes al conocimiento de Doru lineare (Dermaptera, Forficulidae), un insecto de importancia agronómica en el Noroeste argentino. CIRPON Revista de Investigación X (1-4): 13-18. ISSN 0326-3789.

Murua, M. G., P. Fidalgo and E. Virla. 2003. First record of Trichogramma atopovirilia Otman & Platner (Hym., Trichogrammatidae) for Argentina, attacking the eggs of Spodoptera frugiperda Smith (Lep., Noctuidae) on corn. Vedalia (México) 9-10: 25-26. ISSN 1405-0420.

Olmi, M and E. Virla. 2006. Familia Dryinidae. Capitulo 35, 401-418. In: Fernández, F. and M. Sharkey (eds). Introducción a los Hymenoptera de la Región Neotropical. Sociedad Colombiana de Entomología y Univ. Nac. de Colombia., Bogota D.C., xxx + 894 pp. ISBN 958-701-708-0.

Olmi, M. and E. G. Virla. 2008. A new species of Anteon Jurine (Hymenoptera: Dryinidae) from Argentina. Florida Entomologist 91: 159-161. ISSN 0015-4040.

Olmi, M. and E. Virla. 1993. Contribution to the knowledge of the Dryinidae of Argentina (Hymenoptera, Chrysidoidea). Phytophaga (Palermo, Italia) 4 (1993): 57-67. ISSN 0393-8131.

Olmi, M. and E. Virla. 2004. Description of two new species of Dryinidae (Hymenoptera: Chrysidoidea) from Argentina. Zootaxa 709: 1-7. ISSN 1175-5326, 1175-5334 (on line).

Olmi, M., E. Virla and F. Fernández C. 2000. Las Avispas Dryinidae de la Región Neotropical (Hymenoptera: Chrysidoidea). Biota Colombiana 1(2): 141-163. ISSN 0124-5376. 2000.

Remes Lenicov, A. M., Olmi M. and E. Virla. 2009. Phrictopyga holmgreni (Muir): New Record for Argentina and Description of a New Parasitoid, Gonatopus concinnus (Hemiptera: Delphacidae et Hymenoptera: Dryinidae). Journal of the Kansas Entomological Society 82(2): 174-182. ISSN 0022-8567.

Remes Lenicov, A.M.M. de and E. Virla. 1993. Homópteros auquenorrincos asociados al cultivo de trigo en la República Argentina. I. Análisis preliminar de la importancia relativa de las especies. Studies on Neotropical Fauna and Environment (Ecology and Systematics)(Alemania), 28 (4): 211-222. ISSN 0165-0521.

Remes Lenicov, A.M.M. de and E. Virla. 1996. Descripción de los estados inmaduros de Delphacodes haywardi y notas acerca de su comportamiento en condiciones de laboratorio (Homoptera - Delphacidae). Revista de la Sociedad Entomológica Argentina 55 (1-4): 165-174. ISSN 0373-5680.

Remes Lenicov, A.M.M. de and E. Virla. 1999 (2001). Delfácidos asociados al cultivo de maíz en la República Argentina (Insecta - Homoptera - Delphacidae). Revista de la Facultad de Agronomía, La Plata, 104 (1): 1-15. ISSN 0041-8678.

Remes Lenicov, A.M.M. de and E. Virla. 1999. Homopteros vectores de interés fitosanitario: un problema creciente en la Argentina. Revista de la Sociedad Entomológica Argentina 58 (1-2): 34-37. ISSN 0373-5680.

Remes Lenicov, A.M.M. de, E. Virla and G. Varela. 1997. Descripción de los estados inmaduros de Toya propinqua (Fieber) y notas acerca de su comportamiento en condiciones experimentales (Insecta: Homoptera: Delphacidae). Neotropica 43 (109-110): 85-91. ISSN 0548-1686.

Remes Lenicov, A.M.M. de, S. Paradell and E. Virla. 2005. “Homoptera, Auchenorrhyncha”. en Cordo, H., G. Logarzo, K. Braun and O. Di Dorio (Directores). Catálogo de los insectos fitófagos de la Argentina y sus plantas asociadas. 1ra edición, Soc. Entomol. Argentina ediciones, Buenos Aires, Argentina. Pags: 284 (Hom.: Aethalionidae); 330-342 (Hom.: Cercopidae, Cicadellidae); 352-355 (Hom.: Delphacidae); 377-378 (Hom.: Membracidae). ISBN 987-21319-1-0.

Sosa, A.J. , A.M. Marino de Remes Lenicov, R. Mariani, and H.A. Cordo. 2005. Life History of Megamelus scutellaris with Description of Immature Stages (Hemiptera: Delphacidae). Annals of the Entomological Society of America 98(1): 66-72.

Triapitsyn, S. and E. Virla. 2004. A new Anagrus (Hymenoptera: Mymaridae) from Argentina, egg parasitoid of Delphacodes sitarea (Hemiptera: Archaeorrhyncha: Delphacidae). Florida Entomologist 87 (3): 383-385. ISSN 0015-4040.

Triapitsyn, S. V. 1997. The genus Anagrus (Hymenoptera: Mymaridae) in America south of the United States: a review. Ceiba 38 (1): 1-12.

Triapitsyn, S. V. and Virla, E. G. 2004. A new Anagrus (Hymenoptera: Mymaridae) from Argentina, an egg parasitoid of Delphacodes sitarea (Hemiptera: Archaeorrhyncha: Delphacidae). Florida Entomologist 87 (3): 383 - 385.

Triapitsyn, S. V., Logarzo, G. A. and Virla E. G. 2011. Egg Parasitoid of Saccharosydne subandina (Hemiptera: Delphacidae) in Neuquén, Argentina. INTA, Boletín MIP, vol. 20. (http://www.inta.gov.ar/imyza/info/bol/mip/10/bol20/mip20.htm#8)

Virla E. G. and M. Olmi. 2008. Dryinidae. Pp. 357-372. In: Debandi G, Claps LE, Roig-Juñent S. (Eds.), Biodiversidad de Artrópodos Argentinos. Vol II. SEA Ediciones Especiale. ISBN 978-987-21319-2-0.

Virla E. G. and M. Olmi. 2008. Description of two new species of Gonatopus Ljungh from Argentina (Hym., Chrysidoidea, Dryinidae). Journal of the Kansas Entomological Society 81(3): 182-187. ISSN 0022-8567.

Virla E. G., A. Álvarez, F. Loto, L. M. Pera and M. Baigorí. 2008. Fall armyworm strains (Lepidoptera: Noctuidae) in Argentina, their associate host plants and response to different mortality factors in laboratory. Florida Entomologist 91(1): 63-69. ISSN 0015-4040.

Virla E. G., Moya-Raygoza, G. and J. A. Rafael. 2009. First Record of Eudorylas schreiteri (Shannon) (Diptera: Pipunculidae), as a Parasitoid of the Corn Leafhopper (Hemiptera: Cicadellidae) in Argentina, with a Table of Pipunculid-Host Associations in the Neotropical Region. Neotropical Entomology 38(1): 152-154. ISSN: 1519-566X.

Virla, E. 1992 (1995). Biología de Pseudogonatopus chilensis Olmi, 1989 (Hym.- Dryinidae). Acta Entomológica Chilena 19: 123-127. ISSN 0716-5072.

Virla, E. 1995. Aspects of the biology of Gonatopus desantisi (Hym.- Dryinidae). Frustula Entomologica n.s. (Italia) 17 (30): 29-34. ISSN 0532-7679.

Virla, E. 1997. New Neotropical species of Dryinidae (Hymenoptera - Chrysidoidea). Frustula entomológica (Pisa), n.s. XX (XXXIII): 1-16. ISSN 0532-7679.

Virla, E. 1997. New species of Gonatopodinae from the Neotropics (Hymenoptera: Dryinidae)â**. Bollettino della Societa Entomologica Italiana, Genova, 129 (2): 171-186. ISSN 0373-3491.

Virla, E. 2000. Los ejemplares de Dryinidae (Hymenoptera: Chrysidoidea) de la colección entomológica del Instituto Fundación M. Lillo. Revista de la Sociedad Entomológica Argentina 59 (1-4): 161-170. ISSN 0373-5680.

Virla, E. 2001. Inedit Data on Neotropic Gonatopodinae, with descriptions of a gynader specimen (Hymenoptera, Chrysidoidea). Revista de la Sociedad Entomológica Argentina 60 (1-4): 81-88. ISSN 0373-5680.

Virla, E. 2002 (2004) Biología de Gonatopus bonaerensis (Hymenoptera: Drynidae), enemigo natural de Delphacidae (Hemiptera) en Argentina. Revista de la Facultad de Agronomía, La Plata, 105 (2): 18-26. ISSN 0041-8678.

Virla, E. 2003. Evidences of induced maternal molt inhibition by Gonatopus chilensis (Insecta, Hym.: Dryinidae) in Peregrinus maidis (Insecta, Hem.: Delphacidae) nymphsâ**. Neotropica 49: 35-41. ISSN 0548-1686.

Virla, E. 2004. Biología de Anagrus flaveolus (Hymenoptera, Mymaridae), parasitoide del vector del “Mal de Rio Cuarto del maíz”, Delphacodes kuscheli (Hemiptera, Delphacidae). Acta Zool. Lilloana 47(1-2): 137-148. ISSN 0065-1729.

Virla, E. and J. A. Rafael. 1997. Datos bionómicos preliminares y descripción de la hembra de Cephalops penepauculus (Hardy) (Diptera - Pipunculidae) un parasitoide de Delphacidae (Auchenorrhyncha) en Argentina. CIRPON Revista de Investigación X (1-4): 33-36. ISSN 0326-3789.

Virla, E. and M. Olmi. 1998. Presencia de Haplogonatopus hernandezae Olmi, 1984 (Hymenoptera, Dryinidae) en Argentina. Acta Zoologica Lilloana 44 (2): 409-410. ISSN 0065-1729.

Virla, E. and M. Olmi. 1993 (1995). Description of Gonatopus virlai, new species from Argentina, and first data on its development stages (Hymenoptera, Dryinidae). Fragmenta Entomologica (Italia), XXVI (1): 85-94. ISSN 0429-288X.

Virla, E. and M. Olmi. 1998. The Dryinidae of Argentina (Hymenoptera - Chrysidoidea). Acta Entomológica Chilena 22: 19-35. ISSN 0716-5072.

Virla, E. and M. Olmi. 2001. Los ejemplares de Dryinidae (Insecta - Hymenoptera) depositados en la colección del Museo de La Plata. Neotropica 47: 61-80. ISSN 0548-1686.

Virla, E. and René Maragliano. 1993. Preferencias alimentarias y sitios de oviposición de Delphacodes haywardi (Muir, 1929) en diferentes hospedadores, en condiciones de laboratorio (Insecta - Homoptera - Delphacidae). Revista de la Sociedad Entomológica Argentina 52 (1-4): 101-106. ISSN 0373-5680.

Virla, E. and S. Mangione. 2000. Morfología de los estados preimaginales de Gonatopus chilensis, y consideraciones sobre las estructuras relacionadas a la nutrición de sus larvas inmaduras (Hymenoptera: Dryinidae). Neotropica 46: 37-49. ISSN 0548-1686.

Virla, E. G. 2003. Un nuevo huésped para Cheiloneurus cupreicollis (Hym.: Encyrtidae), parasitoide de Dryinidae (Hym.) en Argentina. Revista de la Sociedad Entomológica Argentina 62(1-2): 63-64. ISSN 0373-5680.

Virla, E. G. and M. Olmi. 2007. New records of Dryinidae from Argentina and descriptions of two new species (Hymenoptera: Chrysidoidea). Bollettino di Zoologia agraria e di Bachicoltura, ser. ii, 39 (3): 165-184. ISSN 0336-2403.

 

---

 

Below is a now dated compilation of specific records of predators and parasites of delphacids.  Initially, it was intended to find all such records and compile them, but it was fairly quickly discovered that there are a very large number of reports of predators and parasites of delphacids (for the parasites, see table above) .  This table is just the beginning of a compilation of records of delphacid predators and parasites, and will be periodically updated.

 

A list of predators and parasites recorded for delphacid planthoppers.
Note:  a list of predators and parasites specific to Nilaparvata lugens can be found through the CAB International, 2005. Crop Protection Compendium, but also see link above to ricehoppers.net..
Predator family	Predator Genus	Predator Species	Planthopper Prey	Stage Attacked	Host Plant	Locality	Reference
"Spiders"			Prokelisia marginata, Nilaparvata lugens, Sogatella furcifera, Prokelisia marginata, Dicranotropis, Peregrinus maidis, Javesella pellucida, Stenocranus minutus	nymph, adult	Spartina alterniflora (USA), Oryza satvia (SE Asia, Sri Lanka, Malaysia, Philippines), Holcus mollis (UK), Zea mays (Hawaii), Cereal crops, Dactylis glomerata		Vince et. Al (1981), Chiu (1979), Fowler (1987), Ng (1978), Manti (1989), Pfeiffer and Wiegert (1981), Waloff and Thompson (1980), Napompeth (1973), Raatikainen (1967)
Araneidae			Laodelphax striatellus	nymph, adult	Oryza satvia (Japan)		Kobayashi and Hiwada (1968)
Araneidae	various (viz. Argiope bruennichi (Scop.), Araneus quadratus Clerck, Araneus diadematus Clerck, Nuctenea cornuta (Clerck), Meta segmentata Clerck, and Tetragnatha extensa (L.), )		Stenocranus major				Nyffeler and Benz, 1989
Argiopidae	Araneus	inustus	Nilaparvata lugens		Oryza satvia	Taiwan	Chiu et al (unpubl.)
Argiopidae	Neoscona	doenitzi	Nilaparvata lugens		Oryza satvia	Japan	Kayashima (1960)
Carabidae	Acupalpus	inornatus	Nilaparvata lugens		Oryza satvia	Taiwan	Chiu et al (unpubl.)
Carabidae	Bembidion	semilunium	Nilaparvata lugens		Oryza satvia	Taiwan	Chiu et al (unpubl.)
Carabidae	Casnoidea	cyanocephala	Nilaparvata lugens		Oryza satvia	Malaysia	Lim (1974)
Carabidae	Casnoidea	infersititrialis	Nilaparvata lugens		Oryza satvia	Sri Lanka, Malaysia	Lim (1974), Fernando (1975), Otake et al (1976)
Carabidae	Ophionea	indica	Nilaparvata lugens		Oryza satvia		Otake (1976)
Coccinellidae	Brumoides	suturalis	Sogatella furcifera, Peregrinus maidis	nymph, adult	Oryza satvia (India), Sorghum halepense (India),		Garg and Sethi (1983), Fisk et al. (1981)
Coccinellidae	Casnoidea	interstitialis	Nilaparvata lugens	nymph, adult	Oryza satvia (Malaysia)		Lim et al. (1978), Ooi (1988)
Coccinellidae	Ceratomegilla	maculata	Tagosodes orizicolus	nymph, adult	Oryza satvia (Central America)		King and Saunders (1984)
Coccinellidae	Coccinella	arcuata	Nilaparvata lugens	nymph, adult	Oryza satvia (India),		Abraham and Mathew (1975)
Coccinellidae	Coccinella	septempunctata	Laodelphax striatellus, Conomelus anceps	nymph	Oryza satvia (Russia), Juncus effesus (UK)		Harpaz (1972), Rothschild (1966)
Coccinellidae	Coelophora	inaequalis	Perkinsiella saccharicida	nymph	S. officinarum (hawaii)		Swezey (1936)
Coccinellidae	Harmonia	sp.	Nilaparvata lugens		Oryza satvia	Philippines	Dyck and Orlido (unpubl.)
Coccinellidae	Hippodamia	tredacimpunctata	Nilaparvata lugens		Oryza satvia	Mainland China	Lei and Wang (1958)
Coccinellidae	Illeis	indica	Peregrinus maidis	nymph, adult	Sorghum bicolor (India)		Fisk et al. (1981)
Coccinellidae	Micraspis	discolor	Nilaparvata lugens	nymph, adult	Oryza satvia (Thailand, Malaysia)		Yasumatsu et al. (1975)
Coccinellidae	Micraspis	vincta	Nilaparvata lugens	nymph, adult	Oryza satvia (Thailand)		Yasumatsu et al. (1975)
Coccinellidae	Synharmonia	octomaculata	Nilaparvata lugens		Oryza satvia (Malaysia)		Lim et al. (1978), Ooi (1988)
Coccinellidae	Verania	sp.	Nilaparvata lugens		Oryza satvia	Philippines	Dyck and Orlido (unpubl.)
Dryinidae	Echthrodelphax	bicolor	Nilaparvata lugens	nymph, adult	Oryza satvia	Japan	Esaki and Hashimoto (1936)
Dryinidae	Echthrodelphax	fairchildi	Nilaparvata lugens	nymph, adult	Oryza satvia	India	Rai (pers. Comm)
Dryinidae	Haplogonatopus	japonicus	Nilaparvata lugens	nymph, adult	Oryza satvia	Japan	Esaki and Hashimoto (1936), Sakai (1932), Esaki and Mochizuki (1941)
Dryinidae	Haplogonatopus	sp.	Nilaparvata lugens	nymph, adult	Oryza satvia	India, sri Lanka	Rai (pers. Comm), Santa et al (unpubl.)
Dryinidae	Pseudogonatopus	flavifemur	Nilaparvata lugens	nymph, adult	Oryza satvia	Japan	Esaki and Hashimoto (1933, 1936), Esaki (1932), Sakai (1932)
Dryinidae	Pseudogonatopus	hospes	Nilaparvata lugens	nymph, adult	Oryza satvia	Thailand	Napompeth (unpubl.)
Dryinidae	Gonatopus	bonaerensis Virla	Delphacodes sitarea Remes Lenicov et Tesón, D. latus Remes; Delphacodes kuscheli Fennah, Metadelphax propinqua (Fieber), Peregrinus maidis (Ashmead) and Dicranotropis sp., Lenicov et Tesón		various	Argentina	Virla 2004
Elenchidae	Elenchus	laponicus	Nilaparvata lugens	nymph, adult	Oryza satvia	Japan	Esaki and Hasimoto (1932), Esaki (1932), Sakai (1932), Mochida and Okada (1973), Kuno (1973)
Elenchidae	Elenchus	koebelei	Nilaparvata lugens	nymph, adult	Oryza satvia	Fiji	Hinckley (1963)
Elenchidae	Elenchus	yasumatsui	Nilaparvata lugens	nymph, adult	Oryza satvia	Thailand	Kifune and Hirashima (1975), Yasumatsu et al (1975), Otake (1976)
Elenchidae	Elenchus	sp.	Nilaparvata lugens	nymph, adult	Oryza satvia	Sri Lanka	Santa et al (unpubl.)
Encyrtidae	Chrysopophagus	australiae	Nilaparvata lugens	nymph, adult	Oryza satvia	Solomon Island	MacQuillan (1974)
Encyrtidae	Echthrogonatopus	exitiosus	Nilaparvata lugens	nymph, adult	Oryza satvia	Solomon Island	MacQuillan (1974)
Eulophidae	Ootetrastichus	nr beatus	Nilaparvata lugens	egg	Oryza satvia	Fiji	Hinckley (1963)
Eulophidae	Tetrastichus	sp.	Nilaparvata lugens		Oryza satvia
Formicidae	Tetramorium	guineense	Nilaparvata lugens		Oryza satvia	Taiwan	Fukuda (1934)
Halictophagidae	Halictophagus (s.s.)	stenocrani	Stenocranus minutus			Hapan	Kifune, 1986
Linyphiidae	Atypena	formosana	Nilaparvata lugens, Sogatella furcifera	nymph, adult	Oryza satvia (Philippines)		Reddy and Heong (1991)
Linyphiidae	Erigonidium	graminicolum	Nilaparvata lugens, Sogatella furcifera	nymph, adult	Oryza satvia (China)		Chiu (19840
Linyphiidae	Gnathonarium	dentatum	Nilaparvata lugens	nymph, adult	Oryza satvia (Korea)		Lee and Park (1977)
Linyphiidae	Grammonota	inornata	Prokelisia marginata	nymph, adult	Spartina alterniflora (USA)		Vince et. al (1981)
Linyphiidae	Lepthyphantes	tenuis	Javesella pellucida, Conomelus anceps	nymph, adult	Grasses (UK), Juncus effusus (UK)		Waloff (1980), Rothschild (1966)
Linyphiidae	Linyphia	clathrata	Conomelus anceps	nymph, adult	Juncus effusus (UK)		Rothschild (1966)
Linyphiidae	Linyphia	triangularis	Conomelus anceps	nymph, adult	Juncus effusus (UK)		Rothschild (1966)
Linyphiidae	Notioscopus	pallidulus	Nilaparvata		Oryza satvia	Japan	Sakai (1932)
Lycosidae	Lycosa	modesta	Prokelisia dolus; Prokelisia marginata	nymph, adult	Spartina alterniflora (USA)		Döbel (1987)
Lycosidae	Lycosa	pseudoannulata	Nilaparvata lugens		Oryza satvia	Japan, Taiwan, Philippines	Kiritani et al (1972), Chu and Wang (1972, 1973), Sasaba et al (1970, 1973), IRRi (1974), Otake (1976)
Lycosidae	Lycosa	pullata	Conomelus anceps	nymph, adult	Juncus effusus (UK)		Rothschild (1966)
Lycosidae	Pagiopalpus	atomarius	Perkinsiella saccharicida	nymph, adult	Saccharum officinarum (USA)		Swezey (1936)
Lycosidae	Pardosa	floridana	Prokelisia marginata	nymph, adult	Spartina alterniflora (USA)		Vince et. Al (1981)
Lycosidae	Pardosa	laura	Sogatella furcifera, Laodelphax striatellus	nymph, adult	Oryza satvia (Japan)		Nakamura (1977)
Lycosidae	Pardosa	littoralis	Delphacodes detecta; Tumidagena minuta; Prokelisia dolus; Prokelisia marginata	nymph, adult	Spartina patens (USA); Spartina alterniflora (USA)		Raupp and Denno (1979); Döbel (1987)
Lycosidae	Pardosa	pseudoannulata	Nilaparvata lugens, Sogatella furcifera	nymph, adult	Oryza satvia (Japan, Philippines, Malaysia, China)		Kiritani et al (1972)
Lycosidae	Pardosa	T-insignita	Nilaparvata lugens, Sogatella furcifera	nymph, adult	Oryza satvia (Korea)		Lee and Park (1977)
Lycosidae	Pirata	subpiraticus	Nilaparvata lugens	nymph, adult	Oryza satvia (Korea)		Lee and Park (1977)
Lycosidae	Pirata	sp.	Nilaparvata lugens	nymph, adult	Oryza satvia (Japan, China)		Kobayashi (1975), China (1984)
Mermithidae	Agamermis	unka	Nilaparvata lugens	nymph, adult	Oryza satvia	Japan	Esaki and Hashimoto (1933), Esaki (1932), Imamura (1932), Kaburaki and Imamura (1932), Kuno (1973)
Mesoveliidae	Mesovelia	vittigera	Nilaparvata lugens	nymph, adult	Oryza satvia (Philippines)		Heong et al. (1990)
Micryphantidae	Oedothorax	insecticeps	Nilaparvata lugens	nymph, adult	Oryza satvia (Philippines)		Kenmore (1980)
Miridae	Cyrtorhinus	fulvus	Tarophagus colocasiae	egg	Colocasia esculenta (Hawaii)		Matsumoto and Nishida (1966)
Miridae	Cyrtorhinus	lividipennis	Nilaparvata lugens, Sogatella furcifera, Sogatella kolophon, Peregrinus maidis	egg, nymph	Oryza satvia (Japan, Philippines, Malysia, India, China, SE Asia, Solomon Is., Fiji, Indonesia), Pasture grass (Fiji), Zea mays (Hawaii)		Suenaga (1963), Dyck and Orlido (1977),  Kuno and Dyck (1985),  and Kalode (1987), Qingcai and Jervis (1988), Chiu (1979), Stapley (1976), Hinckley (1963), Otake and Hokyo (1976), Ooi (1982),  Cook and Perfect (1985), Hinckley (1963), Napompeth (1973)
Miridae	Tytthus	alboornatus	Delphacodes detecta	egg	Distichilis spicata		Tallamy and Denno (1979)
Miridae	Tytthus	chinensis	Perkinsiella vittiensis, Nilaparvata lugens	egg, adult (nilaparvata lugens)	S. officinarum (Hawaii), Oryza satvia	Fiji, Japan, solomon Islands, Hawaii	Swezey (1936), Kobayashi (1961), Hinckley (1963), MacQuillan (1968, 1974), (Stapley (1976)
Miridae	Tytthus	mundulus	Nilaparvata lugens, Perkinsiella saccharicida, Peregrinus maidis	egg, adult (Nilaparvata lugens)	S. officinarum (Hawaii, Australia), Zea mays (Hawaii)	Taiwan, Fiji, Hawaii	Swezey (1936), Osborn (1974), Zimmerman (1948), Chiu and Lung (1975), Hinckley (1963)
Miridae	Tytthus	parviceps	Saccharosydne saccharicida, Perkinsiella saccharicida, Nilaparvata lugens	egg, adult (Nilaparvata lugens)	S. officinarum (Jamaica)	India	Metcalfe (1971), Bull (1981), Pathak and Saha (1976)
Miridae	Tytthus	pygmaeus	Conomelus anceps	egg, nymph	Juncus effusus (UK)		Rothschild (1966)
Miridae	Tytthus	vagus	Prokelisia marginata	egg	Spartina alterniflora (USA)		Denno (1983)
Mymaridae	Anagrus	delicatus	Prokelisia marginata		Spartina alterniflora
Mymaridae	Anagrus	flaveolus	Nilaparvata lugens	egg	Oryza satvia	Japan	Yasumatsu and Watanabe (1965)
Mymaridae	Anagrus	incarnatus	Nilaparvata lugens	egg	Oryza satvia	China
Mymaridae	Anagrus	nr flaveolus	Nilaparvata lugens, Sogatella furcifera, Laodelphax striatellus	egg	Oryza satvia	Japan	Otake
Mymaridae	Anagrus	optabilis	Sogatella furcifera, Nilaparvata lugens, Laodelphax striatellus	egg	Oryza sativa, Secale cereale, Polyphagous; Oryza, Avena, Arrhenatherum, Triticum, Zea, hordeum	Thailand	Yasumatsu et al (1975), Miura et al (1979)
Mymaridae	Anagrus	optalilis	Nilaparvata lugens	egg	Oryza satvia	Thailand	Yasumatsu et al. (1975), Nishida et al (1976)
Mymaridae	Anagrus	sp.	Nilaparvata lugens	egg	Oryza satvia	Taiwan, Japan, Malaysia	Lin (1974), Fukuda (1934), Kuno (1973), Heong (pers. Comm.)
Mymaridae	Gonatocerus	sp.	Nilaparvata lugens	egg	Oryza satvia	Thailand	Yasumatsu et al (1975)
Mymaridae	Lymaenon	sp.	Nilaparvata lugens	egg	Oryza satvia	Taiwan	Lin (1974)
Mymaridae	Mymar	indica	Nilaparvata lugens	egg	Oryza satvia	Taiwan	Lin (1974)
Mymaridae	Mymar	taprobanicum	Nilaparvata lugens	egg	Oryza satvia	Thailand	Yasumatsu et al (1975)
Mymaridae	Polynema	sp.	Nilaparvata lugens	egg	Oryza satvia	Thailand	Yasumatsu et al (1975)
Nabidae	Dolichonabis	limbatus	Conomelus anceps	nymph, adult	Juncus effusus (UK)		Rothschild (1966)
Nabidae	Nabis	capsiformis	Laodelphax striatellus,  Perkinsiella saccharicida	nymph, egg	Zea mays (Israel), S. officinarum (Hawaii)		Harpaz (1972), Swezey (1936)
Nabidae	Nabis	flavo marginatus	Conomelus anceps	nymph, adult	Juncus effusus (UK)		Rothschild (1966)
Nabidae	Nabis	sp.	Nilaparvata lugens		Oryza satvia	Japan	Kuno (1973)
Phalangidae	Leiobunum	blackwalli	Conomelus anceps	nymph, adult	Juncus effusus (UK)		Rothschild (1966)
Phalangidae	Oligolophus	agrestis	Conomelus anceps	nymph, adult	Juncus effusus (UK)		Rothschild (1966)
pipunculid fly			Stenocranus minutus	adult	Dactylis glomerata		Calvert and Wilson (1986)
Pipunculidae	Dorylas	sp.	Nilaparvata lugens	nymph, adult	Oryza satvia	Sri Lanka	Santa et al (unpubl.)
Pipunculidae	Pipunculus	javanensis	Nilaparvata lugens	nymph, adult	Oryza satvia	taiwan	Chiui et al (unpubl)
Pipunculidae	Tomosvaryella	oryzaetora	Nilaparvata lugens	nymph, adult	Oryza satvia	Taiwan	Chiui et al (unpubl)
Pipunculidae	Tomosvaryella	epichalca	Nilaparvata lugens	nymph, adult	Oryza satvia	Taiwan	Chiui et al (unpubl)
Pipunculidae	Tomosvaryella	subvirescens	Nilaparvata lugens	nymph, adult	Oryza satvia	Taiwan	Chiui et al (unpubl)
Pipunculidae	Cephalops	curtinfrons	Stenocranus minutus			England	May, 1979
Salticidae			Perkinsiella saccharicida	nymph, adult	S. officinarum (Mauritius)		Williams (1957)
Salticidae	Plexippus	paykulli	Nilaparvata lugens		Oryza satvia	Japan	Sakai (1932)
Salticidae	Icius	magister	Nilaparvata lugens		Oryza satvia	Japan	Sakai (1932)
Salticidae	Zygoballus	bettini	Stenocranus lautus	nymph	Carex lurida		Calvert and Wilson (1986)
Staphylinidae	Paederus	fuscipes	Nilaparvata lugens	nymph, adult	Oryza satvia (Malaysia)		Lim et al. (1978), Ooi (1988)
Staphylinidae	Stenus	cicindelloides	Nilaparvata lugens		Oryza satvia	Taiwan	Chiu et al (unpubl.)
Tetragnathidae	Tetragnatha	laboriosa	Prokelisia marginata	nymph, adult	Spartina alterniflora (USA)		Throckmorton (1989)
Tetragnathidae	Tetragnatha	mandibulata	Prokelisia saccharicida, Nilaparvata lugens	nymph, adult	saccharum officinarum (USA), Oryza satvia (Philippines)		Swezey (1936), Barrion et al. (1981)
Tetragnathidae	Tetragnatha	sp.	Nilaparvata lugens	nymph, adult	Oryza satvia (SE Asia)		Ôtake and Hokyo (1976)
Theridiidae	Enoploynatha	japonica	Nilaparvata lugens		Oryza satvia	Japan	Kiritani et al (1972), Kuno (1973), Otake (1976)
Theridiidae	Theridion	spp.	Nilaparvata lugens		Oryza satvia	Philippines	IRRI (1972)
Theridiidae	Theridion	octomaculatum	Nilaparvata lugens		Oryza satvia	Taiwan	Chiu et al (unpubl.)
Trichogrammatidae	Aphelinoidea	sp.	Nilaparvata lugens	egg	Oryza satvia	Taiwan	Fukuda (1934)
Trichogrammatidae	Oligosita	sp.	Nilaparvata lugens	egg	Oryza satvia	Thailand	Yasumatsu et al (1975)
Trichogrammatidae	Paracentrobia	andoi	Nilaparvata lugens,	egg	Oryza satvia	Taiwan, Japan	Suenaga (1963), Lin (1974)
Trichogrammatidae	Paracentrobia	garuda	Nilaparvata lugens	egg	Oryza satvia	Thailand	Yasumatsu et al (1975)
Trichogrammatidae	Paracentrobia	yasumatsui	Nilaparvata lugens	egg	Oryza satvia	Thailand	Yasumatsu et al (1975)
Trichogrammatidae	Trichogramma	sp.	Nilaparvata lugens	egg	Oryza satvia	Taiwan	Fukuda (1934)
Veliidae	Microvelia	douglasi (including ssp. atrolineata)	Nilaparvata lugens	nymph, adult	Oryza satvia (Philippines, SE Asia)		Cook and Perfect (1985), Chiu (1979)
Veliidae	Microvelia	sp.	Nilaparvata lugens	nymph, adult	Oryza satvia (Malaysia)		Ooi (1982)